- Preface
- Introduction
- Part One Main Features of Structure and Function
- 1 Structure of the Neuron and Organization of Nervous Tissue
- 2 Glia
- 3 Neuronal Excitability
- 4 Synaptic Function
- 5 Neurotransmitters and Their Receptors
- 6 Parts of the Nervous System
- 7 The Coverings of the Brain and the Ventricular System
- 8 The Blood Supply of the CNS
- Part Two Development, Aging, and Plasticity
- 9 Prenatal and Postnatal Development
- 10 The Nervous System and Aging
- 11 Restitution of Function after Brain Damage
- Part Three Sensory Systems
- 12 Sensory Receptors in General
- 13 Peripheral Parts of the Somatosensory System
- 14 Central Parts of the Somatosensory System
- 15 Pain
- 16 The Visual System
- 17 The Auditory System
- 18 The Sense of Equilibrium
- 19 Olfaction and Taste
- Part Four Motor Systems
- 20 Motor Systems and Movements in General
- 21 The Peripheral Motor Neurons and Reflexes
- 22 The Motor Cortical Areas and Descending Pathways
- 23 The Basal Ganglia
- 24 The Cerebellum
- 25 Control of Eye Movements
- Part Five The Brain Stem and the Cranial Nerves
- 26 The Reticular Formation: Premotor Networks, Consciousness, and Sleep
- 27 The Cranial Nerves
- Part Six The Autonomic Nervous System
- 28 Visceral Efferent Neurons: The Sympathetic and Parasympathetic Divisions
- 29 Sensory Visceral Neurons and Visceral Reflexes
- 30 The Central Autonomic System: The Hypothalamus
- Part Seven Limbic Structures
- 31 The Amygdala and Other Neuronal Groups with Relation to Emotions
- 32 The Hippocampal Formation: Learning and Memory
- Part Eight The Cerebral Cortex
- 33 The Cerebral Cortex: Intrinsic Organization and Connections
- 34 Functions of the Neocortex
- Literature
- Index
(p. 557) The Cerebral Cortex
Chapter 33 deals with the anatomic organization of the cerebral cortex at two levels. The first concerns information processing in a small volume of cortex. The second concerns the interconnection of different cortical units with long association and commissural fibers, forming task-specific neural networks. The neurons of the neocortex are arranged in six layers with different properties. Further, the cortical neurons are arranged in smaller modules, often in the form of columns. There are two main kinds of cortical neuron: the pyramidal cells with long axons destined for other parts of the cortex or subcortical targets and interneurons (with numerous subtypes) with short axons remaining in the cortical gray matter. The pyramidal cells are glutamatergic while all interneurons are GABAergic. In addition, modulatory transmitters regulate the cortical excitability level. The cortex is divided into cytoarchitectonic areas that differ with regard to connections and functional specializations.
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- Preface
- Introduction
- Part One Main Features of Structure and Function
- 1 Structure of the Neuron and Organization of Nervous Tissue
- 2 Glia
- 3 Neuronal Excitability
- 4 Synaptic Function
- 5 Neurotransmitters and Their Receptors
- 6 Parts of the Nervous System
- 7 The Coverings of the Brain and the Ventricular System
- 8 The Blood Supply of the CNS
- Part Two Development, Aging, and Plasticity
- 9 Prenatal and Postnatal Development
- 10 The Nervous System and Aging
- 11 Restitution of Function after Brain Damage
- Part Three Sensory Systems
- 12 Sensory Receptors in General
- 13 Peripheral Parts of the Somatosensory System
- 14 Central Parts of the Somatosensory System
- 15 Pain
- 16 The Visual System
- 17 The Auditory System
- 18 The Sense of Equilibrium
- 19 Olfaction and Taste
- Part Four Motor Systems
- 20 Motor Systems and Movements in General
- 21 The Peripheral Motor Neurons and Reflexes
- 22 The Motor Cortical Areas and Descending Pathways
- 23 The Basal Ganglia
- 24 The Cerebellum
- 25 Control of Eye Movements
- Part Five The Brain Stem and the Cranial Nerves
- 26 The Reticular Formation: Premotor Networks, Consciousness, and Sleep
- 27 The Cranial Nerves
- Part Six The Autonomic Nervous System
- 28 Visceral Efferent Neurons: The Sympathetic and Parasympathetic Divisions
- 29 Sensory Visceral Neurons and Visceral Reflexes
- 30 The Central Autonomic System: The Hypothalamus
- Part Seven Limbic Structures
- 31 The Amygdala and Other Neuronal Groups with Relation to Emotions
- 32 The Hippocampal Formation: Learning and Memory
- Part Eight The Cerebral Cortex
- 33 The Cerebral Cortex: Intrinsic Organization and Connections
- 34 Functions of the Neocortex
- Literature
- Index